Nuparu

Nuparu is the Matoran genius who is responsible for inventions like the Vahki and Boxor.

Metru Nui
On Metru Nui, Nuparu invented the Kralhi, but they were too powerful and the Krahli became to be known as "Nuparu's folly". He later invented a better version of the Krahli, the Vahki. One day, he was wandering in the Fikou Web when he discovered protodermis leaking into the Archives. He immediately warned the Toa Metru, and Vakama and his team went off to seal the cracks. Later, he was called to the Coliseum and stuffed into a Matoran Pod by one of his own creations, the Vahki.

Mata Nui
On Mata Nui, in the Gahlok attack on Onu-Koro, Nuparu, Onepu and Taipu were separated from the other Onu-Matoran and trapped. While trapped, Nuparu took apart deactivated Gahlok and built the Boxor, a weapon used to free himself and his companions and later to battle Bohrok.

Later, when all of Le-Koro save Tamaru and Kongu were taken over by krana, Nuparu and Takua helped to free Le-Koro.

When the Matoran traveled back to Metru Nui and Jaller was gathering a band of Matoran to go to Voya Nui, Nuparu was the Onu-Matoran Jaller chose.

Voya Nui
Nuparu, along with Jaller and the other Matoran, traveled to Karzahni, where Nuparu's orange Pakari was replaced with a black Kadin, and from there traveled to Voya Nui each one in a Toa Canister. These canisters were struck by lightning from the Red Star upon arrival to Voya Nui. When Nuparu emerged, he was a Toa, specifically a Toa of Earth.

As a Toa, Nuparu controlled earth and had lightning mixed in with his earth powers. He wore the Kadin, Mask of Flight.